Tapsoba had an extra-time goal ruled out for offside against Zakaria Sanogo, and Gabon keeper Amonome denied Tapsoba and Ouedraogo in the second added period to send the game to penalties.

It was a fourth win in a row for Reading – a club record in the WSL for the Royals, who move level on points with Manchester City, while Birmingham remain two points adrift of Leicester at the bottom of the table.

Jess Park missed a glorious opportunity when she raced through one-on-one in the second half – goalkeeper Manuela Zinsberger making a crucial save from close range, before also denying Hemp and Weir.

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It is Tottenham’s first defeat in the league since Antonio Conte, a Premier League and FA Cup winner while in charge of Chelsea, replaced the sacked Nuno Espirito Santo in November.

It is the 12th time the Seagulls have drawn in the Premier League this season and extends their unbeaten run to six games, with a top-10 finish and challenge for the European places now looking likely.

Vieira and Palace were aggrieved at key moments in the defeat. There was a suggestion Roberto Firmino was offside in the build-up to Oxlade-Chamberlain’s goal and the penalty award for Jota was controversial.

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